Athearn uses plastic couplers
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Athearn uses plastic couplers because they own McHenry. Accurail likewise uses Accumate couplers for obvious reasons
Athearn's McHenry are junk. I once ran a 20 car (out of the box) Thrall gon train at a modular meet which were factory equipped with the older McHenry's; the train would separate randomly because the plastic finger closure spring was no good having fatigued while in the box. That is basically an intrinsic design defect from the factory. Due to it, I've heard Athearn will replace them with the newer version FWIW (not much).
The McHenry's on newer release Athearn models look decent but are also junk. They tend to jam in the open position. I haven't run them that much and had many jams. They are really bad. Shame on Athearn for using them.
Athearn may save money but the McHenry's are horrible and must be replaced. I hate to say it but horn hooks are probably more reliable. I buy Athearn models that fit my needs despite the horrible couplers they come with.
Accurail Accumates seem to have given me less trouble than McHenry but they are the worst looking of the plastic and take more force to couple. I believe Rob has reported operational issues.
Best practice is to toss the plastic clones as soon as possible.
Correct me if I'm wrong, but Atlas has offered some of the former BLMA freight cars with Kadee's; BLMA factory equipped their models with Kadee's.
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The companies who use the Kadees scale head couplers tend to be the ones that offer higher end and detailed (and expensive) cars. They probably have determined that their customers, who are paying a premium price for accurate cars, are going to want the closest to scale coupler from the gold standard for such things.
The fly in that ointment is Athearn Genesis, who offer higher end and detailed (and yes, expensive cars) . The street price is a bit lower than Tangent but higher than Intermountain, both equipped with scale head Kadee's. For some reason Athearn hasn't come to the same conclusion while selling similar higher end and detailed models (Genesis).
Regardless, it's nearly universal that scale head are used in RTR cars factory equipped with Kadee's.
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As for Walthers and ScaleTrains.com (particularly Rivet Counter), who both have recently created their own metal couplers, I can only assume that they both want the cachet of having a "higher end" (metal instead of plastic) coupler, and believe they can make them reliably for cheaper than they can buy them from Kadee.
Walther Protomax being nearly identical clones to the Kadee #5 head seem to operate almost as well and most seem to be ok leaving them installed.
By contrast, lots of modelers report ScaleTrains metal couplers aren't as good as Kadee's and say they replace them. Replacing any plastic coupler, including ST, seems to be a given.
Given the overall cost of models, the cost of Kadee's seems to be a smaller and smaller fraction of the cost of a freight car and especially an engine. It seems the benefit of factory equipping Kadee's would outweigh the small increase in cost to factory equip.
Modelers who already know Kadee are the "gold standard" are going to absorb that cost anyway, one way or the other. Those who don't may "pay" with unreliable beahavior and frustration. Fortunately, we can buy Kadee's and make up for that particular deficiency from Atlas and Athearn, and Scale Trains (assuming their metal couplers are less reliable than Kadee's.
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I've started putting #153's on most everything as they have a shorter shank giving closer, more prototypical coupling. #158's for cars with cushion draft gear that have more of the drawbar exposed.
Do they seem to operate just as well as the standard shank. The shorter shank does seem like they would improve model appearance and over a long train, may shorten the length significantly.